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Susan Golden and her lab members ask how a circadian clock functions at the molecular level as a timekeeping mechanism, and how it is integrated at the cellular level to control gene expression and metabolism. She works with the only developed model organism for circadian rhythms in a prokaryote, the cyanobacterium Synechococcus elongatus, and applies tools of genetics, genomics, biochemistry, and structural biology. Golden developed the BioClock Studio, a course that pairs undergraduate students with graduate and postdoctoral mentors to create publicly available educational videos, graphics, and electronic texts for circadian biology topics.