Dr. Tontonoz is also Professor of Pathology and Laboratory Medicine at the University of California, Los Angeles. He received his undergraduate degree in molecular biology and biochemistry from Wesleyan University and his M.D. and Ph.D. degrees from Harvard Medical School. While earning his Ph.D. degree, he worked with Bruce Spiegelman at the Dana-Farber Cancer Institute. He carried out his postdoctoral training with Ron Evans at the Salk Institute and completed his residency training in pathology at the University of California, San Diego.