FOR FURTHER
INFORMATION:

Jennifer Donovan
(301) 215-8859
donovanj@hhmi.org
Howard Hughes
Medical Institute
4000 Jones Bridge Road Chevy Chase, MD 20815-6789
(301) 215-8500

Ninety-two graduate students and 69 medical students have won new fellowships from the Howard Hughes Medical Institute. The predoctoral fellowships are five-year awards for students working for doctoral degrees in the biological sciences. Medical student fellowships enable medical students to devote a year during their medical schooling to biomedical research and provide support for some to resume their medical education.

Through these two programs, HHMI now supports nearly 500 students at more than 70 universities and research centers. Since the fellowship programs began, the Institute has awarded nearly $200 million to 2,000 graduate and medical students.

This year's predoctoral fellows will study at 27 institutions in the United States. Fellows include 48 men and 44 women. Eight are minorities underrepresented in the sciences. Medical student fellows, including 40 men, 16 women and four from minorities underrepresented in the sciences, will conduct biomedical research at 19 U.S. institutions.

Predoctoral Fellowships in Biological Sciences

2001 Fellows

Cell and Developmental Biology Genetics and Molecular Biology Immunology and Microbiology Mathematical Biology and Epidemiology Neuroscience and Physiology Structural Biology and Biochemistry

Cell and Developmental Biology

Name	Undergraduate Institution	Fellowship Institution
Jennifer J. Bainter	Washington University	Washington University
Anna Greka	Harvard University	Harvard University
Kristina M. Herbert*	University of Pennsylvania	Stanford University
Madhulika Jain	Massachusetts Institute of Technology	University of California, San Francisco
Melissa E. Marks	Grinnell College	Stanford University
David T. Miyamoto	Harvard University	Harvard University
Margot G. Paulick	University of Wisconsin, Madison	University of California, Berkeley
John P. Rearick	Purdue University Main Campus	Harvard University
Madolyn L. Rogers	University of South Florida	Stanford University
Catherine A. Schowe*	Rice University	University of Wisconsin, Madison
Heather L. Stickney	Swarthmore College	Stanford University
Samantha Sutton	University of Illinois at Urbana, Champaign	Massachusetts Institute of Technology
Teresa A. Sweat	Kansas State University	Oregon State University
Michael P. Verzi	College of New Jersey	University of California, San Francisco
* Award deferred fron 2000.

Genetics and Molecular Biology

Name	Undergraduate Institution	Fellowship Institution
Orkun Akin	California Institute of Technology	University of California, San Francisco
Siobain M. Duffy*	Rutgers, The State University of New Jersey	Yale University
Ruth A. Dworak Barkley	Texas A & M University	University of Texas Health Science Center
Jose Garcia-Garcia	University of Havana	Oklahoma State University Main Campus
Nathan C. Geething	Vanderbilt University	Stanford University
Christopher T. Hittinger	Southeast Missouri State University	University of Wisconsin, Madison
Jason R. Kennerdell	University of Pittsburgh, Pittsburgh Campus	University of California, San Francisco
Jacob R. LaPorte	University of Delaware	Harvard University
Suzanne R. Lee	Wellesley College	University of California, Berkeley
Derrick H. Lenz	Drew University	Princeton University
Drew M. Lowery	Penn State University Park	Massachusetts Institute of Technology
Erin K. McKittrick	Carleton College	University of Washington
Ellen M. Mills	University of Notre Dame	Washington University
Sandra S. Murray	University of Massachusetts	Stanford University
Venkataramanan I. Nandagopal	Anna University	University of California, Berkeley
Eric J. Peters	Michigan State University	University of California, San Francisco
Richard L. Possemato	Massachusetts Institute of Technology	Harvard University
Todd A. Sangster	Harvard University	The University of Chicago
Joshua A. Shapiro	Williams College	The University of Chicago
* Award deferred from 2000.

Immunology and Microbiology

Name	Undergraduate Institution	Fellowship Institution
Michelle A. Aaron	Clarion University of Pennsylvania	Yale University
Jane Garrity	California Institute of Technology	University of Wisconsin, Madison
Ryan N. Hagglund	Northwestern University	The University of Chicago
Alison E. Hall	Tulane University	University of Pennsylvania
Raymond J. Lynch	Boston College	Yale University
Adam G. Matthews	Harvard University	Harvard University
Priscilla A. McDowell	Ohio Wesleyan University	University of Wisconsin, Madison
Esther A. Obeng	University of Miami	University of Miami
Alexandra E. Purdy	College of William and Mary	University of California, San Diego
Rebecca R. Saff	Texas A & M University	Boston University
Monte M. Winslow	University of Victoria	Stanford University
Virginia A. Young	Clemson University	Wake Forest University

Mathematical Biology and Epidemiology

Name	Undergraduate Institution	Fellowship Institution
Amy R. Chapin	University of Maryland College Park	The Johns Hopkins University
Derek Y. Chiang	University of North Carolina at Chapel Hill	University of California, Berkeley
Gregory M. Cooper	Miami University	Stanford University
Jonathan A. Gelfond	University of Texas at Austin	University of North Carolina at Chapel Hill
Catherine F. Laine	Swarthmore College	Harvard University
John P. Novembre	Colorado College	University of California, Berkeley
Maya L. Petersen	Stanford University	University of California, Berkeley
Christopher D. Snow	Massachusetts Institute of Technology	Stanford University

Neuroscience and Physiology

Name	Undergraduate Institution	Fellowship Institution
Christopher Brechtel	Harvard University	University of California, San Diego
Yeang H. Ch'ng	Yale University	Harvard University
Marlene R. Cohen	Massachusetts Institute of Technology	Stanford University
Fazila P. Erciyas	University of the Bosphorus	University of California, San Francisco
Robert C. Froemke	Tufts University	University of California, Berkeley
Kassatihun D. Gebre-Amlak	Florida A & M University	Vanderbilt University
Isaac A. Hubner	University of Delaware	Harvard University
Oleg A. Igoshin	Novosibirsk State University	University of California, Berkeley
Pakorn Kanchanawong	Cornell University	Stanford University
Sebastian Kraves	University of Buenos Aires	Harvard University
Boris Krupa	The University of Chicago	Massachusetts Institute of Technology
John A. MacKay	Massachusetts Institute of Technology	University of California, San Francisco
Katharine E. Munckton	University of California, Davis	University of California, Berkeley
Katherine I. Nagel	The University of Chicago	University of California, San Francisco
Robert O'Hagan	Wesleyan University	Columbia University
Christine M. Paczkowski	Duke University	University of California, San Diego
Jessica A. Panzer	Cornell University	University of Pennsylvania
Kacy J. Redd	University of Southern Mississippi	Columbia University
Jill A. Sakai	University of Rochester	University of Wisconsin, Madison
Neville E. Sanjana	Stanford University	Massachusetts Institute of Technology
Alexandra M. Schnoes	University of Wisconsin, Madison	University of California, San Francisco
David Schoppik	New York University	University of California, San Francisco
Alykhan F. Shamji	Harvard University	Harvard University
Douglas L. Sheridan	Pennsylvania State University	Yale University
Merek C. Siu	University of British Columbia	University of California, Berkeley
Ryuji Suzuki	University of Massachusetts Dartmouth	Massachusetts Institute of Technology
Dougal G. Tervo	University of Oxford, Corpus Christi College	Cold Spring Harbor Laboratory
Kevin C. Wang	Stanford University	Harvard University

Structural Biology and Biochemistry

Name	Undergraduate Institution	Fellowship Institution
Emily E. Arias	Colorado College	Harvard University
Jeffrey E. Barrick	California Institute of Technology	Yale University
Rebecca J. Charnas	Yale University	Massachusetts Institute of Technology
Susan E. Crown	Cornell University	University of California, Berkeley
Ian W. Davis	Vanderbilt University	Duke University
Karen K. Kincaid	University of California, Berkeley	University of Colorado at Boulder
Zachary A. Knight	Princeton University	University of California, San Francisco
Peter O. Krutzik	University of California, Irvine	Stanford University
Michelle M. Meyer	Rice University	California Institute of Technology
David Y. Takeda	Pomona College	Harvard University
Eileen Woo	Harvard University	University of California, San Francisco

Research Training Fellowship for Medical Students
2001 Fellows
Initial Awards-First Year of Research

Cell and Developmental Biology Genetics and Molecular Biology Immunology and Microbiology Neuroscience and Physiology Structural Biology and Biochemistry

*denotes medical school affiliation if different from fellowship institution

Cell and Developmental Biology

Name	Fellowship Institution	Department	Mentor
Genevieve A. Ambrose	University of Pittsburgh	Otolaryngology	Jennifer Grandis, M.D., Ph.D.
Anthony J. Apicelli III	Washington University	Neurology	David H. Gutmann, M.D., Ph.D.
Weip Chen	Duke University	Cell Biology	Christopher V. Nicchitta, Ph.D.
James J. Chien	Duke University	Cell Biology	Marc G. Caron, Ph.D.
Gregory J. DeBlasi	Yale University	Orthopedics	Roland Baron, D.D.S., Ph.D.
Eric J. Duncavage	St. Jude Children's Research Hospital	Tumor Cell Biology	Charles J. Sherr, M.D., Ph.D./ Clayton W. Naeve, Ph.D.
Hannah M. Huddleston	Indiana University	Pediatrics	D. Wade Clapp, M.D.
Eric C. Kauffman	The University of Chicago	Surgery	Carrie Rinker-Schaeffer, Ph.D.
Paul J. Kim	Yale University	Pathology	Dario C. Altieri, M.D.
Stephanie Misono	Harvard University	Neurology	Thomas L. Schwarz, Ph.D.
Benjamin J. Moeller	Duke University	Radiation Oncology	Mark W. Dewhirst, D.V.M., Ph.D.
Yvonne Ou	Bringham & Women's Hospital (Harvard University*)	Pathology	Michael A. Gimbrone, Jr., M.D.
Shannon M. Sheu	Stanford University	Epithelial Biology	Paul A. Khavari, M.D., Ph.D.
Kevin M. Sullivan	University of Pittsburgh	Pediatrics	Seth J. Corey, M.D.
Alexi A. Wright	University of California, San Francisco (University of Pennsylvania*)	Stomatology	Susan J. Fisher, Ph.D.
Srinivasan Yegnasubramanian	The Johns Hopkins University	Oncology	William G. Nelson, M.D., Ph.D.

Genetics and Molecular Biology

Name	Fellowship Institution	Department	Mentor
David S. Aaronson	Mount Sinai	Pharmacology	Ravi Iyengar, Ph.D.
Tanya R. Hathaway	University of Washington	Oral Biology	Beverly A. Dale-Crunk, Ph.D./ Richard B. Presland, Ph.D.
Junne Kamihara	Massachusetts Institute of Technology (Harvard University*)	Biology	David E. Housman, Ph.D./ Ann M. Graybiel, Ph.D.
Maxwell S.H. Laurans	Yale University	Genetics	Richard P. Lifton, M.D., Ph.D.
Timothy Y. Tseng	Duke University	Gastroenterology	Don C. Rockey, M.D.
Andrew Z. Wang	Harvard University	Human Genetics	D. Gary Gilliland, M.D., Ph.D.
Laura Y. Young	Duke University	Genetics	Joseph Heitman, M.D., Ph.D.

Immunology and Microbiology

Name	Fellowship Institution	Department	Mentor
William R. Burns	Yale University	Pathology	Jordan S. Pober, M.D., Ph.D.
Andrea A. Chow	Children's Hospital of Philadelphia (University of Pennsylvania*)	Pediatrics	Paul A. Offit, M.D.
Christina E. Clark	Harvard University	Microbiology	Darren E. Higgins, Ph.D.
Shaun M. Honig	Mount Sinai	Gene Therapy and Molecular Medicine	Jonathan S. Bromberg, M.D., Ph.D.
Brian H. Ladle	The Johns Hopkins University	Oncology	Elizabeth M. Jaffee, M.D.
Somnath Mookherjee	University of Washington	Medicine	M. Julie McElrath, M.D., Ph.D.
Ngoc T. Phan	Harvard Medical School	Pathology	Joseph G. Sodroski, M.D.
Jeremy B. Richards	Washington University	Pathology and Immunology	Herbert W. Virgin, IV, M.D., Ph.D.
Harris S. Rose	Harvard University	Surgery	Joren C. Madsen, M.D., Ph.D.
Eric J. Seeley	Stanford University (The University of Chicago*)	Rheumatology and Immunology	C. Garrison Fathman, M.D.
Manish R. Sharma	University of Michigan	Surgery	James J. Mule, Ph.D./ Laura M. Beretta, Ph.D.
Laura E. Stewart	Duke University	Medicine	Dhavalkumar D. Patel, M.D., Ph.D.
Parsia A. Vagefi	Harvard University (Yale University*)	Surgery	David H. Sachs, M.D.

Neuroscience and Physiology

Name	Fellowship Institution	Department	Mentor
Edward F. Chang	University of California, San Francisco	Otolaryngology	Michael M. Merzenich, Ph.D.
Patrick C. Chang	J. David Gladstone Institutes (University of California, San Francisco*)	Neurology	Steven M. Finkbeiner, M.D., Ph.D.
Alice S. Chen	Massachusetts General Hospital (Harvard University*)	Neurology	Jang-Ho Cha, M.D., Ph.D./ Anne Young, M.D., Ph.D.
Ted K. Chuang	University of California, San Francisco	Neurological Surgery	Scott C. Baraban, Ph.D./ Samuel J. Pleasure, M.D., Ph.D.
James L. Frazier	The Johns Hopkins University	Neurosurgery	Henry Brem, M.D.
Daniel Friedman	Case Western Reserve University	Neuroscience	Ben W. Strowbridge, Ph.D.
David S. Grenda	Washington University	Oncology	Daniel C. Link, M.D.
James E. Hansen	University of California, Los Angeles	Physiological Science	Rachelle H. Crosbie, Ph.D.
Asim Haque	Washington University (University of Mississippi*)	Speech and Hearing Sciences	J. David Dickman, Ph. D.
Karen J. Ho	Brinham & Women's Hospital (Harvard University*)	Medicine	James K. Liao, M.D.
Benjamin D. Hoehn	Stanford University	Neurosurgery	Gary K. Steinberg, M.D., Ph.D. Midori A. Yenari, M.D.
James C. Kao	Duke University	Surgery	Henry S. Friedman, M.D.
Michael C. Ke	Stanford University	Neurobiology	Jennifer L. Raymond, Ph.D.
Robert J. Kotloski	Duke University	Neurology	James O. McNamara, M.D.
Samuel D. Madoff	University of Rochester	Neurobiology and Anatomy	M. Kerry O'Banion, M.D., Ph.D. Berislav V. Zlokovic, M.D., Ph.D.
Carlo O. Martinez	Washington University	Neurology	John W. McDonald, M.D., Ph.D.
Nader Sanai	University of California, San Francisco	Neurosurgery	Arturo Alvarez-Buylla, Ph.D.
Tarra M. Wright	Duke University	Radiology	Gregory McCarthy, Ph.D.

Structural Biology and Biochemistry

Name	Fellowship Institution	Department	Mentor
Mittul Gulati	University of California, San Francisco	Physiology	Vishwanath R. Lingappa, M.D., Ph.D.
Ryan B. Turner	Harvard University	Nutrition	Guy L. Reed, M.D.

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