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FOR FURTHER
INFORMATION:


Jennifer Donovan
(301) 215-8859
donovanj@hhmi.org
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Medical Institute
4000 Jones Bridge Road Chevy Chase, MD 20815-6789
(301) 215-8500


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June 29, 1999
New Graduate Program Awards

The Howard Hughes Medical Institute has awarded 165 fellowships to enable some of the world's most talented young scientists to receive advanced training in biomedical research.

Eighty-seven of the fellowships are five-year awards for graduate students pursuing doctoral degrees in the biological sciences. The other 78 fellowships will enable medical students to take a year off from their medical studies to pursue research full time, or to resume their medical studies after a year of research.

"It is gratifying to note the continuing strength of interest in biomedical research among these outstanding graduate students and medical students," said HHMI President Purnell W. Choppin. "As the fellows pursue their training with superb mentors in excellent university and medical school departments, the HHMI awards will help place them firmly on the path to important discoveries in the future."

HHMI, one of the nation's largest philanthropies, now supports 500 students at 74 academic institutions through these two programs. A third fellowship program enables physicians with clinical experience to receive training in basic research for three years. Through fiscal year 1999 HHMI has awarded $200 million in fellowship support to 2,000 outstanding students and physician scientists.

"The fellows have identified challenging and far-reaching areas of research in which to immerse themselves," said Joseph G. Perpich, HHMI's vice president for grants and special programs. "The topics extend across such fundamental issues as the workings of biological clocks and how ion channel structure and function relates to nerve cell actions, as well as the pathogenesis and treatment of devastating health problems such as Alzheimer's disease and cancer. We look forward to learning about their findings, through progress reports and fellows' meetings."

Information about fellows, their research, and the fellowship programs is available on the World Wide Web at www.hhmi.org/fellowships .

Predoctoral Fellowships in Biological Sciences

The Predoctoral Fellowships in Biological Sciences support students for up to five years while they pursue full-time study toward a Ph.D. or Sc.D. degree. The program now supports 393 predoctoral students at an annual cost of more than $12 million.

Each predoctoral award includes annually a $16,000 stipend and a $15,000 cost-of-education allowance, which goes to the fellowship institution. Eligible fields include biochemistry, bioinformatics, biophysics, biostatistics, cell biology, developmental biology, epidemiology, genetics, immunology, mathematical and computational biology, microbiology, molecular biology, neuroscience, pharmacology, physiology, structural biology, and virology. This year's 87 new fellows, including 47 men and 40 women, are planning graduate study at 29 academic institutions in the United States, 2 in the United Kingdom, and 1 in Germany. Six of the fellows are from minority groups underrepresented in the sciences. The group includes 72 U.S. citizens and 4 fellows from Canada, 3 from the People's Republic of China, and 1 each from Argentina, Belgium, Czech Republic, Ethiopia, Japan, Russia, and Singapore.

Research Training Fellowships for Medical Students

The Research Training Fellowships for Medical Students will enable 60 medical students to conduct biomedical research full time in a laboratory of their choice for one year. The 43 men and 17 women selected this year will study at 28 institutions. Six of the fellows are from minority groups underrepresented in the sciences. Each $27,000 award includes a $16,000 stipend, a $5,500 research allowance, and a $5,500 allowance to the fellowship institution.

In addition to these new fellowships to medical students for a research year, HHMI also announced that it would provide continued support to 12 men and 6 women who were fellows during the past year. The 18 continued fellows are returning to school to complete their medical studies and will receive a stipend and education allowance totaling $31,000 each year for up to two years. Later this year, additional awards for continued support will be made to medical students in the HHMI-NIH Research Scholars program, bringing the overall medical student fellowship program to an annual level of more than $3 million.


The Howard Hughes Medical Institute employs scientists in cell biology, genetics, immunology, neuroscience, and structural biology. More than 320 Hughes investigators conduct medical research in HHMI laboratories at 71 outstanding academic medical centers and universities nationwide. Through its complementary grants program, HHMI also supports science education in the United States and a select group of researchers abroad.

Predoctoral Fellowships in Biological Sciences 1999 Fellows

Cell and Developmental Biology

Name

Undergraduate Institution,
Year of Graduation

Fellowship Institution

Francis Joseph Alenghat

Harvard University, 1997

Harvard University

Kirsten Bomblies

University of Pennsylvania, 1996

University of Wisconsin-Madison

Christine Beensen Campbell

Smith College, 1996

Duke University

Maria Eugenia Chiappe

University of Buenos Aires (Argentina), 1997

Rockefeller University

Haley Vinson Hieronymus

Wellesley College, 1999

Harvard University

Richard Goff James

University of Michigan-Ann Arbor, 1998

Harvard University

Elizabeth Christine Marin

University of California-San Diego, 1996

Stanford University

Gabriela Motyckova

Wellesley College, 1996

Harvard University

Tomas Diego Perez

Massachusetts Institute of Technology, 1998

Stanford University

Jason Mark Rihel

West Virginia University, 1998

Harvard University

Adrienne H. K. Roeder

Stanford University, 1999

University of California-San Diego

Serena Jean Silver

New York University, 1999

Massachusetts Institute of Technology

Ann Elizabeth Vernon

University of Tulsa, 1998

University of Cambridge, St. John's College (United Kingdom)



Genetics and Molecular Biology

Name

Undergraduate Institution,
Year of Graduation

Fellowship Institution

Amy Anne Caudy

Washington University, 1999

Cold Spring Harbor Laboratory

Brad Alan Chapman

Michigan State University, 1999

University of Georgia

Jayatri Das

Pennsylvania State University Main Campus, 1999

Princeton University

Christina Hwei-Lin Eng

Massachusetts Institute of Technology, 1998

University of California-Berkeley

Brendan Daniel Galvin

University of California-San Diego, 1996

Massachusetts Institute of Technology

Baruch Ze'ev Harris

Brown University, 1995

University of California-San Francisco

Melissa Marian Harrison

Harvard University, 1998

Massachusetts Institute of Technology

Jerry Yung-Chi Hsu

Harvard University, 1997

Stanford University

Van Anh Thi Huynh

Indiana University at Bloomington, 1998

University of Cambridge, Peterhouse College (United Kingdom)

Ravi Shanker Kamath

Harvard University, 1997

University of Cambridge, Peterhouse College (United Kingdom)

Karen Sue Kim

University of Rochester, 1997

University of California-San Francisco

John Isaac Murray

Carnegie Mellon University, 1999

Stanford University

Anna Theresa Neill

University of Richmond, 1999

University of California-San Diego

Clark James Nelson

University of Wisconsin-Madison, 1990

University of Wisconsin-Madison

Jennifer Danielle Ramond

University of California-Berkeley, 1999

University of California-San Francisco

Wendy Michele Reeves

Kansas State University, 1998

University of Washington

Megan Elizabeth Rokop

Brown University, 1999

Massachusetts Institute of Technology

Jessica Dawn Schiff

University of California-San Diego, 1998

Massachusetts Institute of Technology

Jeff S. Smith

Rice University, 1997

Emory University

Anne Violet Unger

Stanford University, 1998

University of California-San Diego

Mark Samuel Voorhies

University of California-San Diego, 1999

University of California-Berkeley

Amanda Beth Wass

Ohio State University Main Campus, 1999

Harvard University

Nazumi Alice Yamada

Trinity College (Connecticut), 1998

University of North Carolina at Chapel Hill



Immunology and Microbiology

Name

Undergraduate Institution,
Year of Graduation

Fellowship Institution

Nathan Walter Goehring*

Amherst College, 1998

Harvard University

Jennifer Kristin Hartt

University of Maryland College Park, 1999

Harvard University

Brendan Neil Lilley

Princeton University, 1998

Harvard University

Grace Engfang Lim

University of California-Berkeley, 1998

University of California-San Diego

Lubomir Nikolaev Merkov

Whitman College, 1999

Stanford University

Leigh Alice O'Mara

Tufts University, 1998

Washington University

Jeanne Marie Opar

Swarthmore College, 1999

Cornell University

Paul Glyndwr Thomas

Wake Forest University, 1999

Harvard University

Jeremy Michael Yarwood

University of Minnesota-Twin Cities, 1997

University of Minnesota-Twin Cities

*Award deferred from 1998



Mathematical Biology and Epidemiology

Name

Undergraduate Institution,
Year of Graduation

Fellowship Institution

Jef Leo Jose Piet Leroy

State University of Ghent (Belgium), 1998

Cornell University

Mahlet Getachew Tadesse

University of the District of Columbia, 1998

Harvard University

Elizabeth Ellen Thomas

Evergreen State College, 1999

Cold Spring Harbor Laboratory

Lu Tian

Nankai University (People's Republic of China), 1995

Harvard University

Olga Garrievna Troyanskaya

University of Richmond, 1999

Stanford University



Neuroscience and Physiology

Name

Undergraduate Institution,
Year of Graduation

Fellowship Institution

Adam Michael Breier

University of Kentucky, 1999

University of California-Berkeley

Zev David Bryant

University of Washington, 1998

University of California-Berkeley

Jason Patrick Davis

Massachusetts Institute of Technology, 1998

California Institute of Technology

Dion Kai Dickman

Washington University, 1999

Harvard University

Kelly Anne Foster

University of Pittsburgh-Pittsburgh Campus, 1999

Harvard University

Jesse Michael Gray

University of Wisconsin-Madison, 1998

University of California-San Francisco

Christina Gutierrez

Rollins College, 1999

University of Miami

Jason Thomas Huse

Princeton University, 1996

University of Pennsylvania

Elizabeth Ann Kensinger

Harvard University, 1998

Massachusetts Institute of Technology

Timothy Michael Kubow

University of California-Berkeley, 1998

University of California-San Francisco

Mark Kenneth Larson

Concordia College-Moorhead, 1998

University of North Carolina at Chapel Hill

Ji Luo

University of Cambridge (United Kingdom), 1998

Harvard University

Sean Patrick MacEvoy

Brown University, 1997

Brown University

Kurt Wesley Marek

University of California-San Diego, 1994

University of California-San Francisco

Claire Elizabeth McKellar

Yale University, 1997

University of California-Berkeley

Samar Bharat Mehta

Stanford University, 1999

University of California-San Diego

Ann Yukari Nakayama

University of Massachusetts, 1998

Stanford University

Xuan Viet Nguyen

Princeton University, 1998

University of Oklahoma Health Sciences Center

Cristopher Myers Niell

Stanford University, 1995

Stanford University

Liam Michael Paninski

Brown University, 1999

New York University

Zachary Edward Perlman

Carleton College, 1994

Harvard University

Michael Alan Robinson

Brandeis University, 1995

Harvard University

Trent Alan Watkins

University of California-Berkeley, 1997

Stanford University

Leor Weinberger

University of Maryland College Park, 1998

University of California-Berkeley

Anne Marie Wissman

Michigan State University, 1998

University of Washington

Zhou Zhu

Saint Mary's College (Indiana), 1999

Harvard University



Structural Biology and Biochemistry

Name

Undergraduate Institution,
Year of Graduation

Fellowship Institution

Joshua Adam Bittker

Massachusetts Institute of Technology, 1999

Harvard University

Daniel Marc Cohen

Rutgers the State University of New Jersey New Brunswick Campus, 1998

Johns Hopkins University

Thomas Greg Fazzio

University of Utah, 1993

University of Washington

Arvind Govindarajan

University of Toronto (Canada), 1998

Massachusetts Institute of Technology

Christine Diane Hardy

University of California-Los Angeles, 1997

University of California-Berkeley

Daniel Joseph Klein

University of Maryland Baltimore County, 1999

Yale University

Mathew Gregory Miller

University of Oregon, 1997

University of California-San Francisco

Carolyn Marie Ott

University of Nebraska-Lincoln, 1998

University of California-San Francisco

Alexander Ra Pico

University of Oregon, 1998

Rockefeller University

Eliana Saxon

Bryn Mawr College, 1997

University of California-Berkeley

James McKay Turner

California Institute of Technology, 1998

Scripps Research Institute



Research Training Fellowships For Medical Students 1999 Fellows
Initial AwardsFirst Year of Research

Continued Awards

Cell and Developmental Biology

Name

Fellowship Institution

Research Mentor

Department

David Allen Barbie

Harvard University

Edward E. Harlow, Ph.D.

Cancer Center

Rebecca Lynn Brown

University of Pennsylvania

Gregory S. Kopf, Ph.D.

Obstetrics and Gynecology

Kevin Soohyung Choe

Albert Einstein College of Medicine

Arthur I. Skoultchi, Ph.D.

Cell Biology

Caroline Jaeyung Choi

Harvard University

Michael Klagsbrun, Ph.D.

Surgery and Pathology

Anna Lutz

Mount Sinai School of Medicine of the City University of New York

Lynn M. Schnapp, M.D.

Medicine

Sandeep Gopinath Mistry

Baylor College of Medicine

Norman M. Greenberg, Ph.D.

Cell Biology

Drew Moghanaki

University of California-San Diego (Texas Tech University Health Sciences Center School of Medicine*)

Carol L. Macleod, Ph.D.

Medicine

Robert D. Cardiff, M.D., Ph.D.

Pathology

Adarsh Vijay Mudgil

State University of New York at Stony Brook

Jonathan A. Garlick, D.D.S., Ph.D.

Oral Biology and Pathology

Lorne B. Taichman, M.D., Ph.D.

Oral Biology and Pathology

Lily Esther Tang

University of California-San Francisco

Jeff Arbeit, M.D.

Surgery

Jonathan Leif Tueting

University of Chicago

Elaine Fuchs, Ph.D.

Molecular Genetics and Cell Biology

Zhen Wang

Northwestern University

J. Larry Jameson, M.D., Ph.D.

Endocrinology

Kalaiselvi B. Williams

Harvard University (Brown University School of Medicine*)

Marian R. Neutra, Ph.D.

Medicine

Daniel Francis Zlogar

University of California-San Francisco (Duke University School of Medicine*)

Vishwanath R. Lingappa, M.D., Ph.D.

Physiology

* A medical school affiliation other than the fellowship institution is indicated in parentheses.


Genetics and Molecular Biology

Name

Fellowship Institution

Research Mentor

Department

Benjamin Sands Brooke

University of Utah

Mark T. Keating, M.D.

Oncological Science

Dean Y. Li, M.D., Ph.D.

Internal Medicine

Tessa Cigler

Rockefeller University (Duke University School of Medicine*)

Mary Jeanne Kreek, M.D.

Biology of Addictive Diseases

Karl Steven Laforge

Biology of Addictive Diseases

Ryan Robert Davies

Yale University

Richard P. Lifton, M.D., Ph.D.

Molecular Medicine

Ari Justin Green

University of California-San Francisco (Duke University School of Medicine*)

Jorge R. Oskenberg, Ph.D.

Neurology

Stephen L. Hauser, M.D.

Neurology

Terri Joyce Halperin

Duke University

Joseph R. Nevins, Ph.D.

Genetics

Matthew Galen Hartwig

Duke University

Anne Marie Pendergast, Ph.D.

Pharmacology and Cancer Biology

Douglas S. Tyler, M.D.

Surgery

Jeffrey Sean Huo

University of Michigan-Ann Arbor

Kotoku Kurachi, Ph.D.

Human Genetics

Tamiko Robin Katsumoto

University of California-San Francisco

Kevin Mark Shannon, M.D.

Pediatrics

Andy Oliver Miller

Harvard University

Stephen B. Calderwood, M.D.

Infectious Disease

Goutham Narla

Mount Sinai School of Medicine of the City University of New York

Scott Lawrence Friedman, M.D.

Medicine

Barbara Denise Saatkamp

Rockefeller University (Harvard Medical School*)

Jeffrey M. Friedman, M.D., Ph.D.

Genetics

Sue Ann Wee

Stanford University

Paul A. Khavari, M.D., Ph.D.

Dermatology

Brian Matthew Wolpin

Harvard University

David Morse Livingston, M.D.

Cancer Biology

Ningyuan Nancy Xu

Albert Einstein College of Medicine

Nicole B. Schreiber-Agus, Ph.D.

Molecular Genetics

* A medical school affiliation other than the fellowship institution is indicated in parentheses.


Immunology and Microbiology

Name

Fellowship Institution

Research Mentor

Department

Alex S. Carcamo

Harvard University

Thomas S. Kupper, M.D.

Dermatology

Emmanuelle M. Clerisme

Yale University

Jeffrey Bender, M.D.

Cardiovascular Medicine

Megan Anne Cooper

Ohio State University Main Campus

Michael A. Caligiuri, M.D.

Internal Medicine

Kira Rand Giovanielli

Yale University

Joseph E. Craft, M.D.

Rheumatology

Rafaz Hoque

Cornell University Joan and Sanford I. Weill Medical College and Graduate School of Medical Sciences

Roy Silverstein, M.D.

Medicine

Arash Izadpanah

University of California-San Diego

Martin F. Kagnoff, M.D.

Medicine

Gordon Ka Wing Lam

Duke University

Barton F. Haynes, M.D.

Medicine

Alexander Meyer Lesokhin

Albert Einstein College of Medicine

Marshall S. Horowitz, M.D.

Microbiology and Immunology

Celia F. Brosnan, Ph.D.

Pathology

Howard Yeong-Rung Li

Vanderbilt University

Paul Spearman, M.D.

Pediatrics

Miguel Nicolas Rivera

Rockefeller University (Harvard Medical School*)

Ralph M. Steinman, M.D.

Cellular Physiology and Immunology

Roger Takeshi Tomihama

Washington University (University of Hawaii John A. Burns School of Medicine*)

Wayne M. Yokoyama, M.D.

Medicine

Patrick L. Wagner

Tufts University School of Medicine (Harvard Medical School*)

Matthew K. Waldor, M.D., Ph.D.

Medicine

Michael Stuart Waugh

Duke University

Joseph Heitman, M.D., Ph.D.

Genetics

James Andrew Alspaugh, M.D.

Genetics

Alex Keung Wong

Harvard University

Elliot D Kieff, M.D., Ph.D.

Medicine

* A medical school affiliation other than the fellowship institution is indicated in parentheses.


Neuroscience and Physiology

Name

Fellowship Institution

Research Mentor

Department

Carl Joseph Aschkenasi

Harvard University

Joel Keith Elmquist, D.V.M., Ph.D.

Neurology

Jeffrey Flier, M.D.

Medicine

Peter Laurence Castro

Cleveland Clinic Foundation (Vanderbilt University School of Medicine*)

James D. Thomas, M.D.

Cardiology

Isaac S. Cheng

Harvard University (Tufts University School of Medicine*)

Rudy E. Tanzi, Ph.D.

Genetics and Aging

Jen Yuan Chow

Yale University

Joseph Anthony Madri, M.D., Ph.D.

Pathology

Andrew Christopher Cordle

Case Western Reserve University

Gary E. Landreth, Ph.D.

Neuroscience

Christian Paul Erickson

Duke University

Mark Dewhirst, D.V.M., Ph.D.

Radiation Oncology

Karen Friede Holden

University of California-San Francisco

Lennart Mucke, Ph.D.

Neurology

Jean-Christophe Andre Leveque

Duke University

Michael Haglund, M.D., Ph.D.

Neurosurgery

Gregory McCarthy, Ph.D.

Radiology

Darren Lyman Lish

Yale University

Patricia S. Goldman-Rakic, Ph.D.

Neurobiology

Colin Guy Looney

Duke University

James R. Urbaniak, M.D.

Orthopaedic Surgery

Anand Yogesh Mehta

University of California-San Francisco

Robert Haas Edwards, M.D.

Neurology

Bryan Chang-Seok Oh

University of Washington (Stanford University School of Medicine*)

H. Richard Winn, M.D.

Neurological Surgery

Jose J. Otero

Albert Einstein College of Medicine

Jack Kessler, M.D.

Neurology

Albert Jun Pahk

University of Pennsylvania (State University of New York at Stony Brook Health Sciences Center School of Medicine*)

Jeffrey Field, Ph.D.

Pharmacology

Daniel Michael Sciubba

Columbia University

Steven A. Siegelbaum, Ph.D.

Neurobiology

Harvey Edward Smith

Pennsylvania State University Hershey Medical Center

Michael Bruce Smith, Ph.D.

Radiology

Sean Patrick Whalen

University of California-Irvine

Christine M. Gall, Ph.D.

Anatomy and Neurobiology

* A medical school affiliation other than the fellowship institution is indicated in parentheses.


Structural Biology and Biochemistry

Name

Fellowship Institution

Research Mentor

Department

Ravi Kiran Ghanta

Massachusetts Institute of Technology (Harvard Medical School*)

James G. Fujimoto, Ph.D.

Electrical Engineering and Computer Science

Aradhana Mukherjea Venkatesan

Harvard University

Andrea Elizabeth Dunaif, M.D.

Medicine

* A medical school affiliation other than the fellowship institution is indicated in parentheses.


Continued AwardsTwo Years of Support for Completion of Medical Studies

Name

Medical School

Felix Adler

Yale University

Christopher David Barth

Rush University

Andrew William Beckwith

Thomas Jefferson University

Fina CaƱas

Harvard University

Deborah Eva Citrin

Duke University

Abigail Elizabeth Collins

University of California-San Francisco

William Anthony Frisella, Jr.

Washington University

Ankie Marie Hata

Duke University

Aram Francis Hezel

State University of New York at Buffalo

David Kwangmo Hong

Northwestern University

Rajul Krishna Jain

University of Texas Southwestern Medical Center at Dallas

Laura Alice Kirkman

Albert Einstein College of Medicine

Peter Tun-Ho Lin

University of Michigan-Ann Arbor

Adeka Dantani McIntosh

Baylor University

Jason Stephen Ogiste

University of Rochester

Alice Tam

Albert Einstein College of Medicine

Geoffrey Lim Uy

Washington University

Andrew Jenho Yee

Harvard University

   

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