
As vertebrates evolved from aquatic to terrestrial forms, circulation patterns became more complex. The most ancient group, the fish, has an undivided heart that pumps the blood through the gills to the rest of the tissues.

In the youngest group, the birds (but also in mammals), the heart is divided so that one side pumps blood to the lungs (pulmonary circulation) and the other to the rest of the tissues (systemic circulation). This change from an undivided to a divided circulation occurred as a consequence to the change in lifestyle from aquatic to terrestrial habitats.